A reevaluation of the previously described planktonic foraminifera genera and species characteristic for the peri- Carpathian area of the Paratethys suggest the reassignment of the genera Praeorbulina and Velapertina to Candorbulina, with some species introduced into the synonymy of Candorbulina indigena (Luczkowska 1955). The late Badenian (early Serravallian) peri-Carpathian Candorbulina indigena lineage originated in the tropical area of the Indo-Pacific and evolved in the connected parts of the Paratethys until the regional tectonics isolated the basin by the beginning of the Sarmatian (mid Serravallian).
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